Definition

Let — the expected number of offspring per individual, i.e., the mean of the offspring distribution .

  • If : (certain extinction, unless )
  • If : is the smallest positive solution to:

Interpretation

  • (left side): the extinction probability we’re solving for
  • : number of offspring a single individual produces
  • : probability of producing exactly offspring
  • (right side): probability that all offspring lines eventually go extinct (each offspring independently has extinction probability , and there are of them)

The right side sums over all possible offspring counts: probability of having offspring × probability all lineages die out.

This is a self-consistency equation — appears on both sides because extinction of the whole population requires extinction of every sub-lineage.